"Axelrod discovered that when these encounters were repeated over a long period of time with many players, each with different strategies, "greedy" strategies tended to do very poorly in the long run while more "altruistic" strategies did better, as judged purely by self-interest. He used this to show a possible mechanism to explain what had previously been a difficult hole in Darwinian theory: how can apparently altruistic behavior evolve from the purely selfish mechanisms of natural selection? "