http://tolweb.org/tree?group=Primates&contgroup=...

It must not be forgotten that although a high standard of morality gives but a slight or no advantage to each individual man and his children over the other men of the same tribe, yet that an increase in the number of well-endowed men and an advancement in the standard of morality will certainly give an immense advantage to one tribe over another. A tribe including many members who, from possessing in a high degree the spirit of patriotism, fidelity, obedience, courage, and sympathy, were always ready to aid one another, and to sacrifice themselves for the common good, would be victorious over most other tribes; and this would be natural selection.
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It is, however, very difficult to form any judgment why one particular tribe and not another has been successful and has risen in the scale of civilisation.
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Although civilisation thus checks in many ways the action of natural selection, it apparently favours the better development of the body, by means of good food and the freedom from occasional hardships. This may be inferred from civilised men having been found, wherever compared, to be physically stronger than savages.
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With civilised nations, as far as an advanced standard of morality, and an increased number of fairly good men are concerned, natural selection apparently effects but little; though the fundamental social instincts were originally thus gained.